Faculty profile headshot photo of Kathy Martin

Kathy Martin

Senior Research Scientist, Environment and Climate Change Canada

Forest Sciences Centre 3041
2424 Main Mall
Vancouver, BC V6T 1Z4

work phone: 604-822-9695

Areas of research include:

  • Ecology, life history variation, behaviour and conservation of birds in high elevation habitats
  • Metapopulation and connectivity processes for alpine bird populations
  • Avian use of mountain habitats during migration
  • Nest Webs – cavity nesting bird community dynamics in natural and managed forests in relation to forest cutting regimes and natural disturbance events
  • The roles of excavating birds in cavity nesting community dynamics
  • Effectiveness of avian monitoring programs, surrogates and indicator species


Ecology and Life History of Mountain Birds


Life history variation of vertebrates across elevations


Effects of forest management and forest health on avian biodiversity of mixed forest ecosystems in Canada and Chile

Environment Canada

Structure and function of cavity-nester communities in the Americas

NSERC and Environment Canada


Doris Huestis Speirs Award, Outstanding Lifetime Contributions to Canadian Ornithology, Canada 2008
5NR Science Award to Leaders in Sustainable Development, Natural Resources, Canada 2003
Ian McTaggart-Cowan Lifetime Achievement Award, The Wildlife Society, Canadian Section 2016
Godman Salvin Prize, Distinguished Ornithological Work and Contributions to the Ornithological Community, British Ornithologists’ Union 2018

Selected Publications

Altamirano, T. A., M. T. Honorato, J.T. Ibarra, M. de la Maza, D. R. de Zwaan, C. Bonacic, and K. Martin. (2019). Elevation has contrasting effects on avian and mammal nest traits in temperate mountains. Austral Ecology, doi:10.1111/aec.12718
Fuglei, E, J-A Henden, R. Brittas, C. Callahan, J. Hansen, R. A. Ims, A. P. Isaev, J. Lang, C. McIntyre, R. Merizon, O. Y. Mineev, Y. N. Mineev, D. Mossop, O. Nielsen, E. Nilsen, Å. Ø. Pedersen, N. M. Schmidt, B. Sittler, M. Hørnell Willebrand, T. Willebrand, and K. Martin. (2019). Circumpolar status of arctic ptarmigan: Population dynamics and trends. Special report in AMBIO, invited.
De Zwaan, D. R., A.F. Camfield, E. C. MacDonald, and K. Martin (2019). Variation in offspring development is driven more by weather and maternal condition rather than predation risk. Functional Ecology, DOI: 10.1111/1365-2435.13273
Cockle, K., M, K. Trczinski, K. L. Wiebe, A. B. Edworthy, and K. Martin. (2019). Lifetime productivity of tree cavities used by cavity-nesting vertebrates in temperate and subtropical forests. Ecological Applications
De Zwaan, D. R. , S. A. Trefry, and K. Martin. (2018). Efficiency and fitness consequences of two trapping methods for the re-capture of ground-nesting songbirds. Journal of Field Ornithology, 89: 363-371. DOI: 10.1111/jofo.12270.
Scridel, D., M. Brambilla, K. Martin, A. Lehikoinen, A. Iemma, M. Anderle, S. Jahnig, E. Caprio, G. Bogliani, P. Pedrini, A. Rolando, R. Arlettaez, and D. E. Chamberlain A. Rolando, R. Arlettaez, and D. E. Chamberlain. (2018). A review and meta-analysis of the effects of climate change on Holarctic mountain and upland bird populations. Ibis (invited). 160: 489-515. DOI: 10.1111/ibi.12585.
Langin, K.M., C.L. Aldridge, J.A. Fike, R.S. Cornman, K. Martin, G.T. Wann, A.E. Seglund, M.A. Schroeder, C.E. Braun, D.P. Benson, B.C. Fedy, J.R. Young, S. Wilson, D.H. Wolfe, and S.J. Oyler-McCance. (2018). Characterizing range-wide divergence in an alpine-endemic bird: a comparison of genetic and genomic approaches. Conservation Genetics, 19: 1471-1485.
Norris, AR, K. A. Aitken, K. Martin, and S. Pokorny. (2018). Nest boxes increase reproductive output for Tree Swallows in a forest grassland matrix in central British Columbia. PLOS ONE, 13(10): e0204226.
Goullaud, E.L., D.R. de Zwaan, and K. Martin. (2018). Predation risk-induced adjustments in provisioning behavior for horned larks (Eremophila alpestris) in British Columbia. Wilson Journal of Ornithology, 130: 180-190.
Drake, A. and K. Martin. (2018). Local temperatures predict breeding phenology but do not result in breeding synchrony among a community of resident cavity-nesting birds. Scientific Reports, 8:2756. DOI:10.1038/s41598-018- 20977-y
de Zwaan, D. R. and K. Martin. (2018). Substrate and structure of ground nests have fitness consequences for an alpine songbird. Ibis, 160: 790-804. DOI 10.1111/ibi.12582
Edworthy, A.B., M.K. Trzcinski, K.L. Cockle, K.L. Wiebe and K. Martin. (2018). Tree cavity occupancy by nesting vertebrates across cavity age. Journal of Wildlife Management, 82: 639-648. DOI:10.1002/jwmg.21398
St-Amand, J, J. A. Tremblay, and K. Martin (2018). Foraging ecology of the Williamson’s sapsucker: implications for forest management. Condor, 120: 680-702. DOI:10.1650/CONDOR-17-238.1
van der Hoek, Y. and K. Martin (2018). Woodpecker cavity-tree selection in the Ecuadorean Amazon region. Ornitologia Neotropical, 29: 95-99.
Wesolowski, T. and K. Martin. (2018). Tree holes and hole-nesting birds in European and North American Forests. Chapter 4, pp 79-134. In: Ecology and Conservation of Forest Birds. Mikusinski, G, J-M Roberge, and R.J. Fuller, Eds. Ecology, Biodiversity and Conservation Series, Cambridge Univ Press, UK.
Cockle, K. L., K. Martin and A. Bodrati. (2017). Persistence and loss of tree cavities used by birds in the subtropical Atlantic Forest. Forest Ecology and Management, 384: 200-207.
de Zwaan, D.R., J. L. Greenwood and K. Martin. (2017). Feather melanin and micro-structure variation in Dark-eyed Junco (Junco hyemalis) across an elevational gradient in the Selkirk Mountains. Journal of Avian Biology, 48:552-562. DOI: 10.1111/jav.01050.
Altamirano, T. A., J. T. Ibarra, F. Novoa, A. Vermehren, K. Martin, and C. Bonacic. (2017). Roosting records in tree cavities by a forest-dwelling bat species (Histiotus magellanicus) in Andean temperate ecosystems of southern Chile. Revista Bosque, 38 (2): 421-425. DOI: 10.4067/S0717-92002017000200020
van der Hoek, Y., G.V. Gaona, and K. Martin. (2017). The diversity, distribution and conservation status of the tree-cavity nesting birds of the world. Diversity and Distributions, 23:1120-1131. DOI: 10.1111/ddi.12601.
Ibarra, J. T., N. Gálvez, T. A. Altamirano, J. Caviedes, I. M. Rojas, C. Bonacic, and K. Martin. (2017). Seasonal dynamics of avian guilds inside and outside core protected areas in an Andean Biosphere Reserve of southern Chile. Bird Study, Ringing and Migration, 64: 410-420.
Ibarra, J. T., T.A. Altamirano, A. Vermehren, F. Hernán Vargas, and K. Martin. (2017). Observations of a tree-cavity nest of the rufous–legged owl and predation of an owl nestling by a chimango caracara in Andean temperate forests. J. Raptor Research, 51: 85-88. doi/full/10.3356/JRR-16-68.1
Martin, K., S. D. Wilson, E.C. MacDonald, A. F. Camfield, M. Martin, and S. A. Trefrey. (2017). Effects of severe weather on reproduction for sympatric songbirds in an alpine environment: interactions of climate extremes influence nesting success. The Auk, 134: 696-709. DOI:10.1642/AUK-16-271.1
Altamirano, T.A., J. T. Ibarra, K. Martin and C. Bonacic. (2017). The conservation value of tree decay processes as a key driver structuring tree cavity nest webs in South American temperate rainforests. Biodiversity and Conservation, 26:2453-2472. DOI: 10.1007/s10531-017-1369-x.
Ibarra, J. T., M. Martin, K. L. Cockle, and K. Martin. (2017). Maintaining ecosystem resilience: functional responses of tree cavity nesters to logging in temperate forests of the Americas. Scientific Reports, 7:4467. 10.1038/s41598-017-04733-2 (SREP-16-46763-T).
Honorato, M. T., T. A. Altamirano, J. T. Ibarra, M. De la Maza, C. Bonacic, and K. Martin. (2016). Composition and preference for nesting materials of cavity nester vertebrates in temperate Andean forests of Chile. Revista Bosque [in Spanish], 37: 485-492. DOI: 10.4067/S0717-92002016000300005.
Beckmann, C, and K. Martin. (2016). Testing hypotheses about the function of repeated nest abandonment as a life history strategy in a passerine bird. Ibis, 158: 335-342, DOI: 10.1111/ibi.12361
Jackson, M. M., E. Topp, S. E. Gergel, K. Martin, F. Pirotti, T. Sitzia. (2016). Expansion of subalpine woody vegetation over 40 years on Vancouver Island, British Columbia, Canada. Canadian Journal of Forest Research, 46: 437-443, DOI 10.1139/cjfr-2015-0186
Huang A. C., J. E. Elliott, S. Hindmarch, S. Lee, F. Maisonneuve, V. Bowes, K. M. Cheng, and K. Martin. (2016). Increased rodenticide exposure rate and risk of toxicosis in barn owls (Tyto alba) from southwestern Canada and linkage with demographic but not genetic factors. Ecotoxicology, 25: 1061-1071. DOI 10.1007/s10646-016-1662-6
Honorato, M. T., T. A. Altamirano, J. T. Ibarra, M. De la Maza, C. Bonacic, and K. Martin (2016). Composition and preference for nesting materials of cavity nester vertebrates in temperate Andean forests of Chile. Revista Bosque [in Spanish]. 37 (3): 485-492. DOI: 10.4067/S0717-92002016000300005
Hunter, M. Jr., P. Barton, A. Calhoun, J. Pierson, A. Tulloch, M. Westgate, M. Beger, C. Branquinho, T. Caro, J. Gross, J.Heino, P. Lane, C. Longo, K. Martin, W. H. McDowell, C. Mellin, H. Salo, and D. Lindenmayer. (2016). Two roles for ecological surrogacy: indicator surrogates and management surrogates. Ecological Indicators, 63: 121-125.
Seneviratne, S. S., P. Davidson, K. Martin and D. E. Irwin. (2016). Low levels of hybridization across two contact zones among three species of woodpeckers (Sphyrapicus sapsuckers). Journal of Avian Biology, 47:887-898. DOI 10.1111/jav.00946.
Huang, A., J. E. Elliott, K. M. Cheng, K. Ritland, C. Ritland, S. K. Thomsen, S. Hindmarch, and K. Martin. (2016). Barn owls (Tyto alba) in North America: Phylogeographic structure, connectivity, and genetic diversity. Conservation Genetics, 17: 357-367 DOI 10.1007/s10592-015-0787-0.
Floyd, C., and K. Martin. (2016). Avian Ecosystem Engineers: Birds that excavate cavities. Chapter 10, pp 298-320. In: Why Birds Matter, C. Skekercioglu, D. G. Wenny, C. J. Whelan (editors). University of Chicago Press.
Boyle, W.A., B. K. Sandercock, and K. Martin. (2016). Patterns and drivers of intraspecific variation in avian life history along elevational gradients: a meta-analysis. Biological Reviews, 91: 469-482. DOI: 10.1111/brv.12180
MacDonald, E.C., A. F. Camfield, M. Martin, S. Wilson, and K. Martin. (2016). Nest-site selection and consequences for nest survival among three sympatric songbirds in an alpine environment. Journal of Ornithology, 157: 393-405. DOI 10.1007/s10336-015-1286-2.
Drever, M.C., L.W. Gyug, J. Nielsen, A.K. Stuart-Smith, I.P. Ohanjanian and K. Martin. (2015). Using field data and GIS-derived variables to model occurrence of Williamson’s Sapsucker nesting habitat at multiple spatial scales. PLOS ONE, DOI:10.1371/journal.pone.0130849. 10(7): e0130849.
Martin, K., L. A. Robb, S. D. Wilson and C. E. Braun. (2015). White-tailed Ptarmigan (Lagopus leucura). In the Birds of North America Online (A. Poole, Ed.). Ithaca: Cornell Lab of Ornithology; http://bna.birds.cornell.edu/bna/species/068. DOI:10.2173/bna.68
Cockle, K. L. and K. Martin. (2015). Temporal dynamics of a commensal network of cavity-nesting vertebrates: increased diversity during an insect outbreak. Ecology, 96: 1093-1104. http://dx.doi.org/10.1890/14-1256.1
Cockle, K. C., A. Bodrati, M. Lammertink, and K. Martin. (2015). Cavity characteristics, but not habitat, influence nest survival of cavity-nesting birds along a gradient of human impact in the subtropical Atlantic forest. Biological Conservation, 184:193-200. DOI: 10.1016/j.biocon.2015.01.026
Jackson, M., S. E. Gergel and K. Martin. (2015). Citizen science and field survey observations provide comparable results for mapping Vancouver Island White-tailed Ptarmigan (Lagopus leucura saxatilis) distribution. Biological Conservation, 181: 162-172. DOI:10.1016/j.biocon.2014.11.010
Martin, K. J. T., Ibarra, and M. Drever. (2015). Avian surrogates in terrestrial ecosystems: theory and practice. Chapter 5, pp 33-44. In: Indicators and Surrogates in Ecology, Biodiversity and Environmental Management, Edited by D. Lindenmayer, P. Barton and J. Pierson. CSIRO Publishing, Melbourne. CRC Press, London.
Ibarra, J. T., T. A. Altamirano, G. Vergara, F. H. Vargas, and K. Martin. (2015). Natural history and autecology of the Austral Pygmy-Owl (Glaucidium nana) in the Andean temperate forest of the Araucania region, southern Chile. In Edicion Especial: Aves Rapaces Del Bosque Templado Austral [In Spanish]. Boletín Chileno de Ornitología, 21: 29−40.
Ibarra, J. T., and K. Martin. (2015). Beyond species richness: an empirical test of top predators as surrogates for functional diversity and endemism. Ecosphere, 6 (8): 142. http://dx.doi.org/10.1890/ES15-00207.1
Beckmann, C., P. A. Biro, and K. Martin. (2015). Hierarchical analysis of avian re-nesting behavior: mean, across-individual, and intra-individual responses. Behavioral Ecology and Sociobiology, 69:1631–1638. DOI 10.1007/s00265-015-1974-1
Boyle, W.A., and K. Martin. (2015). The conservation value of high elevation habitats to North American migrant birds. Biological Conservation, 192: 461-476. http://dx.doi.org/10.1016/j.biocon.2015.10.008
Jackson, M. M., S. E. Gergel, and K. Martin. (2015). Effects of climate change on habitat availability and configuration for an endemic coastal bird. PLOS ONE, 10(11): e0142110. doi:10.1371/journal. pone.0142110
Ibarra, J. T., and K. Martin. (2015). Biotic homogenization: loss of avian functional richness and habitat specialists in disturbed Andean temperate forests. Biological Conservation, 192: 418-427. http://dx.doi.org/10.1016/j.biocon.2015.11.008
Lindenmayer, D, J. Pierson, P. Barton, M. Beger, C. Branquinho, A. Calhoun, T. Caro, H. Greig, J. Gross, J. Heino, M. Hunter, P. Lane, C. Longo, K. Martin, W. H. McDowell, C. Mellin, H. Salo, A. Tulloch, and M. Westgate. (2015). A new framework for selecting environmental surrogates. Total Science of the Environment, 538: 1029-1038. DOI: 10.1016/ j.scitotenv. 2015.08.056
Martin, K. (2015). Nest webs and woodpecker ecological services: the role of woodpeckers in tree cavity-using wildlife communities in North America. In: Woodpeckers – New Insights Into Their Biology, H. Winkler, ed. Biologie Zentrum, Denisia 36, zugleich Kataloge des oberösterreichischen Landesmuseums Neue Serie 164: 77-86
Edworthy, A. B. and K. Martin. (2014). Long-term dynamics of the characteristics of tree cavities used for nesting by vertebrates. Forest Ecology and Management, 334: 122-128. http://dx.doi.org/10.1016/j.foreco.2014.09.001
Ibarra, J. T., K. Martin, M. C. Drever, and G. Vergara. (2014). Occurrence patterns and niche relationships of sympatric owls in South American temperate forests: a multi-scale approach. Forest Ecology and Management, 331: 281-291. http: dx.doi.org/10.1016/j.foreco.2014.08.027.
Martin, K. (2014). Avian Strategies for living at high elevation: life-history variation and coping mechanisms in mountain habitats. IBIS-Conference Proceeding. http//www. bou.org.uk/bouproc-net/uplands/martin.pdf
MacDonald, E.C., A. F. Camfield, J. E. Jankowski, and K. Martin. (2014). An alpine-breeding songbird can adjust dawn incubation rhythms to annual thermal regimes. The Auk, 131:495-506. DOI: 10.1642/AUK-13-234.1
Ibarra, J. T., K. Martin , T. A. Altamirano, F. H. Vargas, and C. Bonacic. (2014). Factors associated with the detectability of Owls in South–American Temperate Forests: Implications for Nocturnal Raptor Monitoring. Journal of Wildlife Management, 78:1078-1086.
Norris, A.R., and K. Martin. (2014). Direct and indirect effects of an insect outbreak increase the reproductive output for an avian insectivore and nest-cavity excavator, the red-breasted nuthatch (Sitta canadensis). Journal of Avian Biology, 45:280-290. DOI: 10.1111/jav.00112
Krebs, C.J., R. Boonstra, S. Boutin, A.R.E. Sinclair, J.N.M. Smith, B.S. Gilbert, K. Martin, M. O’Donoghue, and R. Turkington. (2014). Trophic Dynamics of the Boreal Forests of the Kluane Region. Arctic, 67: Supplement 71-81. http://dx.doi.org/10.14430/arctic.2012.12-109
Filotas, E., L. Parrott, P. J. Burton, R. L. Chazdon, K. D. Coates, L. Coll, S. Haeussler, K. Martin, S. Nocentini, K. L. Puettmann, F. E. Putz, S. W. Simard, and C. Messier. (2014). Viewing forests through the lens of complex systems science. Ecosphere, 5:art1, pp 1-23. http://dx.doi.org/10.1890/ES13-00182.1
Robles, H and K. Martin. (2014). Habitat-mediated variation in the importance of ecosystem engineers for secondary cavity nesters in a nest web. PLOS ONE, 9(2): e90071. DOI: 10.1371/journal.pone.0090071
Ibarra, J. T., T. A. Altamirano, K. Martin, F. H. Vargas, and C. Bonacic. (2014). Tree cavity-nesting of Austral Pygmy Owls (Glaucidium nana) in Andean Temperate Forests of southern Chile. Journal Raptor Research, 48: 82-85. http://dx.doi.org/10.3356/JRR-13-00037.1
Robles, H and K. Martin. (2013). Resource quantity and quality determine the inter-specific associations between ecosystem engineers and resource users in a cavity-nest web. PLOS ONE, 8(9): e74694. DOI:10.1371/journal.pone.0074694
K. Martin. (2013). The Ecological Values of Mountain Environments and Wildlife. Bentham Publishers. DOI: 10.2174/9781608054886113010004. / Chapter 1, pp 3-29. In: The Impacts of Skiing and Related Winter Recreational Activities on Mountain Environments,
Edworthy, A.B., and K. Martin. (2013). Persistence of tree cavities used by cavity-nesting vertebrates declines in harvested forests. Journal of Wildlife Management, 77: 770-776. DOI: 10.1002/jwmg.526
MacDonald, E.C., A.F. Camfield, J.E. Jankowski, and K. Martin. (2013). Extended incubation recesses by alpine-breeding Horned Larks: a strategy for dealing with inclement weather? Journal of Field Ornithology, 84: 58-68. DOI: 10.1111/jofo.12006
Norris, A.R., M.C. Drever, and K. Martin. (2013). Insect outbreaks increase populations and facilitate reproduction in a cavity-dependent songbird, the Mountain Chickadee (Poecile gambeli). Ibis, 155: 165-176. DOI: 10.1111/ibi.12005
Simard, S.W., Martin, K., Vyse, A., and Larson, B. (2013). Meta-networks of fungi, fauna and flora as agents of complex adaptive systems: an example from the interior Douglas-fir forests of southern British Columbia. Chapter 7, pp 133-164. in: Messier, C., Puettmann, K., and Coates, K.D., Managing Forests as Complex Adaptive Systems: Building resilience to the challenge of Global Change. Earthscan, Taylor & Francis Group, London (Routledge).
Evans Ogden, L. J., K. Martin, and T. D. Williams. (2013). Elevational differences in estimated fattening rates suggest high-elevation sites are high quality habitats for fall migration. The Auk, 130:98-106. http://dx.doi.org/10.1525/auk.2012.11229
Koch, A.J., K. Martin, and K.E.H. Aitken. (2012). The relationship between introduced European Starlings and the reproductive activities of Mountain Bluebirds and Tree Swallows in British Columbia, Canada. Ibis,154: 590-600. DOI: 10.1111/j.1474-919X.2012.01242.x
Blanc, L, and K. Martin. (2012). Identifying suitable woodpecker nest trees using decay selection profiles in trembling aspen (Populus tremuloides). Forest Ecology and Management, 286: 192-202. DOI: 10.1016/j.foreco.2012.08.021
Evans Ogden, L.J., M. Martin, and K. Martin. (2012). Mating and breeding success decline with elevation for the Pacific wren (Troglodytes pacificus) in coastal mountain forests. Wilson Journal of Ornithology, 124: 270-276. DOI: http://dx.doi.org/10.1676/11-186.1
Edworthy, A.B., K.L. Wiebe and K. Martin. (2012). Survival analysis of a critical resource for cavity-nesting communities: patterns of tree cavity longevity. Ecological Applications, 22: 1733-1742. http://dx.doi.org/10.1890/11-1594.1
Wilson, S., and K. Martin. (2012). Influence of life history strategies on sensitivity, population growth and response to climate for sympatric alpine birds. BMC Ecology,12:9. DOI:10.1186/1472-6785-12-9
Norris, A.R., and K. Martin. (2012). Red-breasted nuthatches (Sitta canadensis) increase cavity excavation in response to a mountain pine beetle (Dendroctonus ponderosae) outbreak. Ecoscience, 19: 308-315. DOI: 10.2980/19-4-3519
Cockle, K., K. Martin, and G. Robledo. (2012). Linking fungi, trees, and hole-using birds in a Neotropical tree-cavity network: pathways of cavity production and implications for conservation. Forest Ecology and Management, 264: 210-219. DOI: 10.1016/j.foreco.2011.10.015
Zhu, X., D. Srivastava, J.N.M. Smith, and K. Martin (2012). Habitat selection and reproductive success of Lewis Woodpecker (Melanerpes lewis) at its northern limit. PLOS ONE, 7 (9): e44346, pp. 1-9. DOI: 10.1371/journal.pone.0044346
Aitken, K.E.H., and K. Martin. (2012). Experimental test of nest-site limitation in mature mixed forests of central British Columbia, Canada. Journal of Wildlife Management, 76: 557-565. DOI:10.1002/jwmg.286
Edworthy, A., M.C. Drever, and K. Martin. (2011). Woodpeckers increase in abundance but maintain fecundity in response to an outbreak of mountain pine bark beetles. Forest Ecology and Management, 261: 203-210. DOI:10.1016/j.foreco.2010.10.006
Wilson, S., and K. Martin. (2011). Life history and demographic variation in an alpine specialist at the latitudinal extremes of the range. Population Ecology, 53: 459–471. DOI: 10.1007/S10144-011-0261-X
Cockle, K., K. Martin,and K. Wiebe. (2011). Selection of nest trees by cavity-nesting birds in the Neotropical Atlantic Forest. Biotropica, 43: 228-236. DOI: 10.1111/j.1744-7429.2010.00661.x.
Martin, K., S. Wilson and S. J. Hannon. (2011). Mechanisms underlying variation in renesting ability of willow ptarmigan. Pp. 233–246 in B. K. Sandercock, K. Martin, and G. Segelbacher (editors). Ecology, conservation, and management of grouse. Studies in Avian Biology (no. 39). University of California Press, Berkeley, CA.
Martin, K., and S. Wilson. (2011). Ptarmigan in North America: Influence of life history and environmental conditions on population persistence. in R. T. Watson, T. J. Cade, M. Fuller, G. Hunt, and E. Potapov (Eds.). Gyrfalcons and ptarmigan in a changing world. Vol 1, pp 45-54. The Peregrine Fund, Boise, Idaho, USA. DOI 10.4080/gpcw.2011.0105
Sandercock, B.K., K. Martin, and G. Segelbacher (eds.) (2011). Ecology, Conservation and Management of Grouse. Studies in Avian Biology, No. 39. University of California Press, Berkeley, CA.
Fedy, B.C., and K. Martin. (2011). The influence of fine-scale habitat features on regional variation in population performance of alpine white-tailed ptarmigan. Condor, 113: 306-315.
Koch, A., M.C. Drever, and K. Martin. (2011). The efficacy of common species as indicators; avian responses to disturbance in British Columbia, Canada. Biodiversity and Conservation, 20: 3555-3575. DOI: 10.1007/s10531-011-0148-3
Cockle, K.L., K. Martin, and T. Wesolowski. (2011). Woodpeckers, decay and the future of cavity-nesting vertebrate communities worldwide. Frontiers in Ecology and the Environment, 9: 377-382, DOI: 10:1890/110013
Camfield, A.F., S.F. Pearson, and K. Martin. (2011). A demographic model to evaluate population declines in endangered Streaked Horned Lark. Avian Conservation and Ecology, 6 (2): 4.
K. Martin, Wilson, S., S.J. Hannon. (2011). Mechanisms underlying variation in renesting ability in willow ptarmigan. Studies in Avian Biology, 39: 233-246.
Norris, A.R., K.L. Cockle, and K. Martin (2010). Evidence for tolerance of parasitism in a tropical cavity-nesting bird, the planalto woodcreeper (Dendrocolaptes platyrostris), in northern Argentina. Journal of Tropical Ecology, 26: 619-626.
Cockle, K., M.C. Drever, and K. Martin. (2010). Supply of tree-holes limits nest density of cavity-nesting birds in primary and logged subtropical Atlantic Forest. Biological Conservation, 143: 2851-2857.
Wilson, S., and K. Martin. (2010). Variable reproductive effort for two sympatric ptarmigan in response to spring weather conditions in a northern alpine ecosystem. Journal of Avian Biology, 41:319-326. DOI: 10.1111/j.1600-048X.2009.04945.x
Camfield, A .F., S. F. Pearson, and K. Martin. (2010). Life history variation between high and low elevation subspecies of horned larks (Eremophila spp.). Journal of Avian Biology, 41: 273-281. DOI:10.1111/j.1600-048X.2009.04816.x
Norris, A.R., and K. Martin. (2010). The perils of plasticity: Dual resource pulses increase facilitation but destabilize populations of small cavity-nesting vertebrates. Oikos, 119: 1126-1135. DOI: 10.1111/j.1600-0706.2009.18122.x
Drever, M.C.,and K. Martin. (2010). Response of woodpeckers to changes in forest health and harvest: implications for conservation of avian biodiversity. Forest Ecology Management, 259: 958-966. DOI:10.1016/j.foreco.2009.11.038
Wong, M.L., B.C. Fedy, S. Wilson, and K. Martin. (2009). Adoption in Rock and White-tailed Ptarmigan. The Wilson Journal of Ornithology, 121: 638-641.
Drever, M.C., J. R. Goheen, and K. Martin. (2009). Species-energy theory, pulsed resources, and the regulation of avian richness during a mountain pine beetle outbreak. Ecology, 90:1095-1105.
Holmes, D.J. and K. Martin. (2009). A bird's eye view of aging: What's in it for Ornithologists? Auk, 126:1-23.
Bears, H., K. Martin, and G.C. White. (2009). Breeding in high-elevation habitat results in shift to slower life-history strategy within a single species. Journal of Animal Ecology, 78:365-375.
Camfield, A .F., and K. Martin. (2009). The influence of ambient temperature on horned lark incubation behaviour in an alpine environment. Behaviour, 146:1615-1633.
Martin, M., A.F. Camfield, and K. Martin. (2009). Demography of an alpine population of Savannah Sparrows. Journal of Field Ornithology, 80: 253-264.
Nouvet, S., S. Wilson and K. Martin. (2008). Confirmed breeding of four shorebird species, the Surfbird, Upland Sandpiper, American Golden Plover, and Wandering Tattler, in the Ruby Mountains of the southwest Yukon Territory Western Birds, 39: 22-30.
Mahon, C.L., J.D. Steventon, and K. Martin. (2008). Cavity and bark nesting bird responses to partial cutting in northern conifer forests. Forest Ecology and Management, 265: 2145-2153.
Rangel-Salazar, J.L., K. Martin, P. Marshall, and R.W. Elner. (2008). Habitat variation, nest-site selection and parental behaviour influences on breeding success of the Ruddy-capped Nightingale Thrush (Catharus frantzii): in Chiapas, Mexico. Auk, 125: 358-367.
Mahon, C.L, K. Martin, and V. LeMay. (2008). Do cross-scale correlations confound analysis of nest site selection for chestnut-backed chicadees? Condor, 110: 563-568.
Rangel-Salazar, J.L., K. Martin, P.L. Marshall and R.W. Elner. (2008). Population dynamics of the Ruddy-capped Nightingale Thrush (Catharus frantzii) in Chiapas, Mexico: influences of density, productivity and survival. Journal of Tropical Ecology, 24: 583-593.
Cockle, K., K. Martin, and K. L. Wiebe. (2008). Availability of cavities for nesting birds in the Atlantic Forest, Argentina. Ornitologia Neotropical, 19(Suppl.):269-278.
Aitken, K.E.H., and K. Martin. (2008). Resource selection plasticity and community responses to experimental reduction of a critical resource. Ecology, 89: 971-980.
Fedy, B.C., K. Martin, C. Ritland, and J. Young. (2008). Genetic and ecological data provide incongruent interpretations of population structure and dispersal in naturally subdivided populations of white-tailed ptarmigan (Lagopus leucura). Molecular Ecology, 17: 1905-1917.
Drever, M.C., K.E.H. Aitken, A.R. Norris, and K. Martin. (2008). Woodpeckers as reliable indicators of bird richness, forest health and harvest. Biological Conservation, 141: 624-634.
Cornelius, C., K. Cockle, N. Politi, I. Berkunsky, L. Sandoval, V. Ojeda, L. Rivera, Jr. M. Hunter, and K. Martin. (2008). Cavity-nesting birds in neotropical forests: Cavities as a potentially limiting resource. Ornitologia Neotropical, 19 (Suppl.): 253-268.
Wilson, S. and K. Martin. (2008). Breeding habitat selection of sympatric rock, white-tailed, rock and willow ptarmigan in the southern Yukon Territory, Canada. Journal of Ornithology, 149: 629-637.
Bears, H., M.C. Drever and K. Martin. (2008). Comparative morphology of dark-eyed juncos breeding at two elevations: a common aviary experiment. Journal of Avian Biology, 39: 152-162.
Norris, A. R. and K. Martin. (2008). Mountain pine beetle presence affects nest patch choice of red-breasted nuthatches. Journal of Wildlife Management, 72: 733-737.
Mennill, D.J., T.M. Burg, R.L. Curry, K. Martin, A.R. Norris, L. Ratcliffe, M.W. Reudink, L.M. Rossano, and K.A. Shubert. (2007). Parid reproductive behavior. Chapter 9. Pp. 147-150. In Ken Otter (ed.), Ecology and behavior of chickadees and titmice: An integrated approach. Oxford University Press, UK.
Mahon, C.L., K. Martin, and J.D. Steventon. (2007). Habitat attributes and chestnut-backed chickadee nest site selection in uncut and partial cut forests. Canadian Journal of Forest Research, 37(7): 1272-1285.
Camfield, A.F., A.J. Clason, and K. Martin. (2007). Mother-son parental care in horned larks. Wilson Journal of Ornithology, 199: 302-304.
Drever, M.C. and K. Martin. (2007). Spending time in the forest: responses of cavity nesters to temporal changes in forest health in interior British Columbia. Chapter 13. Pp. 236-251. In J. Bissonnette and I. Storch (eds), Temporal explicitness in landscape ecology; Understanding wildlife responses to changes in time. Springer Verlag, New York.
Martin, K. and A.R. Norris. (2007). Life in the small-bodied cavity nester guild: Demography and reproduction of sympatric Mountain and Black-capped Chickadees within Nest Web communities under changing habitat conditions. Chapter 8. Pp. 111-130. In Ken Otter (ed.), Ecology and behavior of chickadees and titmice: An integrated approach. Oxford University Press. Invited book chapter.
Wiebe, K.L., W.D. Koenig, and K. Martin. (2007). Costs and benefits of nest reuse versus excavation in cavity-nesting birds. Annales Zoologica Fennici, 44: 209-217.
Wilson. S, K. Martin, and S.J. Hannon. (2007). Nest survival patterns in willow ptarmigan: influence of time, nesting stage and female characteristics. Condor, 109:377-388.
Topp, S., L.J. Evans Ogden, and K. Martin. (2007). Apparent absence of blood parasites in winter wrens in British Columbia. Journal of Field Ornithology, 78: 308-313.
Squires, K.A., K. Martin, and R.I. Goudie. (2007). Vigilance behavior in the harlequin duck (Histrionicus histrionicus) during the pre-incubation period in Labrador: Are males vigilant for self or social partner? The Auk, 124:241-252.
Aitken, K.E.H. and K. Martin. (2007). The importance of excavators in hole nesting communities: availability and use of natural tree holes in old and mixed forests of western Canada. Journal of Ornithology, 148 (Suppl. 2): 425-434.
Mahony, N.A., P.G. Krannitz, and K. Martin. (2006). Seasonal fecundity of sagebrush Brewer’s sparrows at the northern edge of the range. The Auk, 123:512-523.
Hannon, S.J. and K. Martin. (2006). Ecology of juvenile grouse during the transition to adulthood Journal of Zoology, 289:422-433. (Invited)
Mahon, C.L. and K. Martin. (2006). Nest survival of chickadees in managed forests: habitat, predator, and year effects. Journal of Wildlife Management, 70:1257-1265.
Martin, K. and K.L. Wiebe. (2006). Impacts of extreme climate events on fitness of alpine birds. Acta Zoologica Sinica, 52 (Supplement):162-164.
Wiebe, K.L., W.D. Koenig, and K. Martin. (2006). Evolution of clutch size in cavity-excavating birds: the nest site limitation hypothesis revisited. American Naturalist, 167:343-353.
Martin, K., A.R. Norris, and M.C. Drever. (2006). Effects of bark beetle outbreaks on avian biodiversity in Interior British Columbia. British Columbia Journal of Ecosystems Management, 7(3):10-24. (Invited paper).
Martin, K. (2005). Aspen: Prime real estate for nesting birds and mammals in Interior Douglas-fir forest in British Columbia Sustainable Forest Management Network advertisement in: Logging and Sawmilling Journal, May, page 51.
Martin, K. (2005). Wildlife biodiversity and MPB [mountain pine beetle] in interior forests. Branchlines, UBC Faculty of Forestry 16(3): 9-10. University of British Columbia, Faculty of Forestry.
Er, K.B.H, J.L. Innes, K. Martin, and B. Klinkenburg. (2005). Forest loss with urbanization predicts bird extirpations in Vancouver. Biological Conservation, 126:410-419.
Sandercock, B.K., K. Martin and S.J. Hannon. (2005). Demographic consequences of age structure in extreme environments: population models for arctic and alpine ptarmigan. Oecologia, 146:13-24.
Wilson, S. and K. Martin. (2005). High elevation habitat associations of birds during fall migration. Ecoscience, 12:561-568.
Robinson, P.A., A.R. Norris and K. Martin. (2005). Interspecific nest sharing by Red-breasted Nuthatch and Mountain Chickadee in British Columbia, Canada. Wilson Bulletin, 117(4): 400-402.
Sandercock, B.K., K. Martin and S.J. Hannon. (2005). Life history variation in extreme environments: comparative demography of arctic and alpine ptarmigan. Ecology, 86: 2176-2186.
Martin, K., G.A. Brown, and J.R. Young. (2004). The historic and current distribution of the Vancouver Island White-tailed Ptarmigan (Lagopus leucurus saxatilis). Journal of Field Ornithology, 75:239-256.
Martin, K. and K. L. Wiebe. (2004). Coping mechanisms of alpine and arctic breeding birds: extreme weather and limitations to reproductive resilience. Integrative and Comparative Biology, 44:177-185.
Leupin, E.E., T.E. Dickinson, and K. Martin. (2004). Resistance of songbirds to habitat perforation in a high elevation conifer forest. Canadian Journal of Forest Research, 34:1919-1928.
Lindh, H. and K. Martin. (2004). Utility of systematic reserve selection approaches for conservation of North American mountain resort communities: a case study in Whistler, Canada. Mountain Research and Development, 24: 319-326.
Aitken, K.E.H. and K. Martin. (2004). Nest cavity availability and selection in aspen-conifer groves in a grassland landscape. Canadian Journal of Forest Research, 34:2099-2109
Melles, S., S. Glenn, and K. Martin. (2003). Urban bird diversity and landscape complexity: Species–environment associations along a multiscale habitat gradient. Conservation Ecology, 7(1): 5. [online] URL: http://www.consecol.org/vol7/iss1/art5
Martin, K. and D.S. Hik. (2003). The State of Ecological and Earth Sciences in Mountain Areas. Science Highlights from symposium on ecological and earth sciences in mountain areas, September 2002, Banff, Alberta, Canada. 46 pages.
Martin, K. (2002). Protocol for monitoring birds in high elevation habitats during fall migration. Unpublished report, Department of Forest Sciences, Univ. of British Columbia and Canadian Wildlife Service, Pacific and Western Region. 3 pages. At: http:// forestry.ubc.ca/alpine/index.htm.
Taylor, L., K. Martin, D.S. Hik and A. Ryall (editors) (2002). Ecological and Earth Sciences in Mountain Areas conference. Banff, AB. The Banff Centre 296 Pages.
Martin, K. (2002). Staying on top: Persistence and connectivity processes for birds in mountain habitats. In: Martin, K. and D.S. Hik (eds), the state of ecological and earth sciences in mountain areas. Science Highlights from symposium on ecological and earth sciences in mountain areas, September 2002, Banff, Alberta, Canada. The Banff Centre. Pp 21-22.
Aitken, K.E.H., K.L. Wiebe, and K. Martin. (2002). Nest-site reuse patterns for a cavity-nesting bird community in interior British Columbia. The Auk, 119: 391-402.
Braun, C. E. and K. Martin. (2001). Unusual summer plumage of white-tailed ptarmigan. Wilson Bulletin, 113: 373-377.
Martin, K. (2001). Wildlife Communities in Alpine and Subalpine Habitats. Chapter 9 pp. 239-260 in D.H. Johnson and T.A. O’Neil (Managing Directors), Wildlife-habitat Relationships in Oregon and Washington. Oregon University Press, Corvallis.
Martin, K., C. Doyle, S.J. Hannon, and F. Mueller. (2001). Herbivores: Forest grouse and Ptarmigan. Chapter 11 pp. 240-260 in C.J. Krebs, S. Boutin and R. Boonstra (eds.), Ecosystem Dynamics of the Boreal Forest: The Kluane Project. Oxford University Press.
Martin, K. and L. Forbes. (2001). Vancouver Island White-tailed Ptarmigan species account. BC Ministry of Environment, Lands and Parks, Victoria, BC. 10p.
Congdon, B.C., J.F. Piatt, K. Martin, and V.L. Friesen. (2000). Mechanisms of population differentiation in marbled murrelets: Historical versus contemporary processes. Evolution, 54:974-986.
Hochachka, W.M., K. Martin, F. Doyle, and C.J. Krebs. (2000). Censusing vertebrate populations using observational data. Canadian Journal of Zoology, 78:521-529.
Martin, K., P.B. Stacey and C.E. Braun. (2000). Recruitment, dispersal, and demographic rescue in spatially-structured white-tailed ptarmigan populations. Condor, 102:503-516. [50]
Wiebe, K.L. and K. Martin. (2000). The use of incubation behaviour by ptarmigan to adjust reproductive investment costs after egg laying. Behavioural Ecology and Sociobiology, 48:463-470.
Steidl, R. J. and L. Thomas (K. Martin, supervisor). (2000). Power analysis and experimental design. In Scneiner, S. M. and J. Gurevitch (eds.) Design and Analysis of Ecological Experiments, 2nd edition. Oxford University Press.
Wiebe, K.L. and K. Martin. (2000). The use of incubation behaviour by ptarmigan to adjust reproductive investment costs after egg laying. Behavioural Ecology and Sociobiology, 48: 463-470.
Martin, K. and J.M. Eadie. (1999). Nest Webs: a community wide approach to the management and conservation of cavity nesting birds. Forest Ecology and Management, 115: 243-25.
Artiss, T., W.M. Hochachka, and K. Martin. (1999). Female foraging and male vigilance in white-tailed ptarmigan (Lagopus leucurus): opportunism or behavioural coordination? Behavioural Ecology and Sociobiology, 46: 429-434.
Boulanger, J., K. Martin, G. Kaiser, and A.E. Derocher. (1999). Evaluating uncertainty in estimating population trends for marbled murrelets. Society of Canadian Ornithologists, Special Pub. No. 1. Biology and Conservation of Forest Birds. pp. 53-63.
Dragon, S., C. Carey, K. Martin, and R. Baumann. (1999). Effect of high altitude and in vivo adenosine/b-adrenergic receptor blockade on ATP and 2,3BPG concentrations in red blood cells of avian embryos. Journal of Experimental Biology, 202: 2787-2795.
Nelson, K.J. and K. Martin. (1999). Thermal aspects of nest-site location for vesper sparrows and horned larks in British Columbia. Studies in Avian Biology, 19:137-143.
Vanderkist, B.A., X.H. Xue, R. Griffiths, K. Martin, W. Beauchamp, and T.D. Williams. (1999). Evidence of male-bias in capture samples of Marbled Murrelets from genetic studies in British Columbia. Condor, 101:398-402.
Easton, W.E. and K. Martin. (1998). The effect of vegetation management on breeding bird communities in British Columbia. Ecological Applications, 48:1092-1103.
Hannon, S.J., P.K. Eason and K. Martin. (1998). Willow Ptarmigan. Pp. 1-28. In: The Birds of North America, No. 369 (A. Poole and F. Gill, Eds.). Philadelphia: The Academy of Natural Sciences; Washington, DC: The American Ornithologists' Union.
Krebs, C.J., A.R.E. Sinclair, R. Boonstra, S. Boutin, K. Martin, and J.N.M. Smith. (1998). Community dynamics of vertebrate herbivores - how can we untangle the web? Chapter 15, pp 447-473. In Herbivores: Between Plants and Predators. H. Olff, V.K. Brown, and R.H. Drent. (editors), Blackwells, Oxford.
Turkington, R.T., E. John, C.J. Krebs, M. Dale, V. Nams, R. Boonstra, S. Boutin, K. Martin, A.R.E. Sinclair and J.N.M. Smith. (1998). The effects of NPK fertilization for nine years on boreal forest vegetation in northwestern Canada. Journal of Vegetation Science, 9:333-346.
Wiebe, K.L. and K. Martin. (1998). Age specific patterns of reproduction in white-tailed (Lagopus leucurus) and willow ptarmigan (L. lagopus). Ibis, 140:14-24.
Wiebe, K.L. and K. Martin. (1998). Costs and benefits of nest cover for ptarmigan: Changes within and among years. Animal Behaviour, 56:1137-1144.
Martin, K. and S. Ogle. (1998). The use of alpine habitats by fall migrating birds on Vancouver Island. Report, Department of Forest Sciences, University of British Columbia and Canadian Wildlife Service, Pacific and Western Region, Delta, B.C. 19 pp. At: http:// forestry.ubc.ca/alpine/index.htm
Martin, K. (1998). The role of animal behavior studies in wildlife science and management. Wildlife Society Bulletin, 26: 911-920.
Carey, C. and K. Martin. (1997). Physiological ecology of incubation of ptarmigan eggs at high and low altitudes. Wildlife Biology, 3:211-218.
Holt, R.F. and K. Martin. (1997). Landscape modification and patch selection: the demography of two secondary cavity-nesting species colonizing clear-cuts. The Auk, 114:443-455.
Wiebe, K.L. and K. Martin. (1997). Effects of predation, body condition and temperature on incubation rhythms of ptarmigan. Wildlife Biology, 3: 218-227.
Hannon, S.J. and K. Martin. (1996). Mate fidelity and divorce in ptarmigan: polygyny avoidance on the tundra. Pp. 192-210. In J.M. Black (ed.), Partnerships in Birds, the Ecology of Long-Term Monogamy. Oxford University Press, London, U.K.
Thomas, L.J. and K. Martin. (1996). The importance of analysis method for the breeding bird survey population trend estimates. Conservation Biology, 10:479-490.
Krebs, C.J., S. Boutin, R. Boonstra, A.R.E. Sinclair, J.N.M. Smith, M.R.T. Dale, K. Martin and R. Turkington. (1995). Impact of food and predation on the snowshoe hare cycle. Science, 269: 1112-1115
Martin, K. (1995). Patterns and mechanisms of age-dependent reproduction and survival in birds. American Zoologist, 35 (4): 123-141.
Martin, K., A. G. Horne, and S. J. Hannon. (1995). The calls and associated behaviour of breeding willow ptarmigan (Lagopus lagopus) in Canada. Wilson Bulletin, 107 (3): 496-509.
Wiebe, K. L., and K. Martin. (1995). Ecological and physiological effects on egg laying intervals in ptarmigan. Condor, 97: 708-717.
S. Boutin, C. J. Krebs, R. Boonstra, M. R. T. Dale, S. J. Hannon, K. Martin, A. R. E. Sinclair, J. N. M. Smith, R. Turkington, M. Blower, A. Byrom, F. I. Doyle, C. Doyle, D. Hik, L. Hofer, A. Hubbs, T. Karels, D. L. Murray, V. Nams, M. O’Donaghue, C. Rohner, and S. Schweiger. (1995). Population changes of the vertebrate community during a snowshoe hare decline in Canada’s boreal forest. Oikos, 74: 69-80.
Artiss, T., and K. Martin. (1995). Male vigilance in paired white-tailed ptarmigan Lagopus leucurus: mate guarding or predator detection? Animal Behaviour, 49: 1249-1258.
Thomas, D., K. Martin, and H. Lapierre. (1994). Doubly-labelled water measures of field metabolic rate in white-tailed ptarmigan: variation in background isotope abundances and effect on CO2 production estimates. Canadian Journal of Zoology, 72: 1967-1972.
Hannon, S. J., K. Martin, L. Thomas, and J. Schieck. (1993). Investigator disturbance and clutch predation in willow ptarmigan: methods for evaluating impact. Journal of Field Ornithology, 64: 575-586.
K. Martin, and A. Horn. (1993). Clutch defense by male and female Willow Ptarmigan (Lagopus lagopus). Ornis Scandinavica, 24:61-266.
K. Martin and C. Wright. (1993). Estradiol cypionate (ECP) markedly improves survival of willow ptarmigan in captivity. Condor, 95:211-217.
C.E. Braun, K. Martin and L.A. Robb. (1993). White-tailed Ptarmigan. In The Birds of North America, No. 68 (A. Poole, P. Stettenheim, and F. Gill, Eds.). Philadelphia: The Academy of Natural Sciences; Washington, DC: The American Ornithologists’ Union. [25]
Martin, K., and D. Hik. (1992). Willow ptarmigan chicks consume moss sporophyte capsules. Journal of Field Ornithology, 63:355-358.
Hannon, S.J., and K. Martin. (1992). Monogamy in willow ptarmigan: is male vigilance important for reproductive success and survival of females? Animal Behavior, 42:747-757.